Wade Bolton Worthen
Rose J. Forgione Professor of Biology
Biology Department
Furman University
Greenville, SC 29613
Office: (864)-294-3390
e-mail: worthen@furman.edu
Courses Taught at Furman University:
1983-88: Graduate Teaching Assistant, Rutgers University
World Dragonfly Association
Dragonfly Society of the Americas
Rose J. Forgione Professor of Biology, Furman University. 2021 - present
Fulbright Scholar, 2023, Dragonfly diversity in Costa Rica
Henry and Ellen Townes Research Professor, Furman University. 1994-1996
Furman Standard Award, Furman University. 2014-2017.
2014-17: “The Effect of Competition and Predation in Dragonfly Communities: Tropic vs. Temperate Comparisons” - FURMAN STANDARD AWARD
2011-12: "Biological Inventory at Furman Forest: Birds, Beetles, and Moths" - $1543
2010-11: "Survey of Dragonflies at State Parks" - $1047
2009-10: "All Taxa Biological Inventory at Furman Forest:
Terrestrial Mammals" -$1260
2008-9: "Competition between dragonfly species: Does size
matter?" - $1088
2005-6: "Morphological and metabolic correlates of resource
use among dragonflies" - $1155
2000-1: "Effect of sediment type on dragonfly larvae" - $680
1999-0: "Classification of larval dragonflies" - $700
1998-9: "Thermal ecology of woodland Drosophila species" -
$595
1993-4: "Interactive effects among the determinants of community
structure" -$500
1991-2: "The effects of competition and predation on the structure
of mycophagous fly communities" -$1000
1990: Knight Foundation, Summer Research Stipend: "Competition,
predation, and the structure of mycophagous fly communities" -$3000
1990-1: "Competition, predation, and the structure of fly communities"
-$650
1989-90: "Coexistence among fly species in model communities"
-$1638
Worthen, W. B., and M. D. Harrell*. 2023. Differences in perch height and response to intruders for territorial and non-territorial Calopteryx maculata (Odonata: Calopterygidae). International Journal of Odonatology, 26: 145–152. link
Worthen, W. B., R. K. Fravel*, and C. P. Horne*. 2021. Downstream changes in odonate (Insecta: Odonata) communities along a suburban to urban gradient: untangling natural and anthropogenic effects. Insects 12(3): 201. link
Worthen, W. B., and M.G. Chamlee*. 2020. Determinants of adult odonate community structure at several spatial scales: effects of habitat type and landscape context. International Journal of Odonatology 23:365-379. link
Rasmussen, P.R., J.A. Self, L. Few, and W. B. Worthen. 2019. Sibling niches and the diagnosis of attention-deficit hyperactivity disorder. The Journal of Individual Psychology, 75:104-121. pdf
Worthen, W. B. 2018. Differences in male-male tandem formation in two species of Micrathyria (Odonata: Libellulidae). Notulae Odonatologicae 9(2) 2018: 60-66.pdf
Worthen, W. B. 2018. Confirming the relationship between body size and perch height in tropical odonates (Odonata: Libellulidae): wet-season contrasts and experimental tests. International Journal of Odonatology pdf
Worthen, W. B., and L. Hoke*. 2018. Pseudoreplication in species comparisons: do individual differences matter? Notulae Odonatologicae 9(1) 2018: 18-25. pdf
Worthen, W. B. 2017. Perch selection in a guild of tropical dragonflies (Odonata: Libellulidae): relationships with body size and thermal ecology. International Journal of Odonatology 20(2): 63-78. DOI: 10.1080/13887890.2017.1330225.
Worthen, W. B. 2016. Observation of wing-whirring behavior in a tropical perching dragonfly, Micrathyria atra (Odonata: Libellulidae). Notulae Odonatologicae, 8(8): 261-265.
Worthen, W. B., and P. H. Morrow*. 2016. Perch selection by three co-occurring species of Celithemis (Odonata: Libellulidae): testing for a competitive hierarchy among similar species. Psyche, 2016: 1-8. Doi: 10.1155/2016/9028105.
Worthen, W. B., and T. M. Hart*. 2016. Resistance to Arrenurus spp. parasitism in odonates: Patterns across species and comparisons between a resistant and susceptible host. Journal of Insect Science 16 (1):1-6. doi: 10.1093/jisesa/iew022
Worthen, W. B., and L. H. Turner*. 2015. The effects of odonate species abundance and diversity on parasitism by water mites (Arrenurus spp.): testing the dilution effect. International Journal of Odonatology 18(3):233-248. DOI: 10.1080/13887890.2015.1081112
Worthen, W. B., and H. J. Horacek*. 2015. The distribution of dragonfly larvae in a South Carolina stream: Relationships with sediment type, body size, and the presence of other larvae. Journal of Insect Science 15: 1-7. DOI: 10.1093/jisesa/iev013
Worthen, W. B., and C. C. Phillips*. 2014. Are community patterns in flight height driven by antagonistic interactions? International Journal of Odonatology 17: 7-26.
Worthen, W.B., and D. Merriman*.2012. Relationships between carabid beetle communities and forest stand parameters: Taxon congruence or habitat association? Southeastern Naturalist 12: 379-386.
Worthen, W. B. 2010. Emergence-site selection by the dragonfly Epitheca spinosa (Hagen). Southeastern Naturalist 9:251-158.
Worthen, W. B. 2010. Flying Dragons: A Colorful field experiment in resource partitioning. American Biology Teacher 72:432-435.
Worthen, W. B. and C. M. Jones*. 2007. The effects of wind speed, competition, and body size on perch height selection in a guild of Libellulidae species (Odonata). International Journal of Odonatology 257-272.
Worthen, W. B., and C. M. Jones*. 2006. Relationships between body size, wing morphology, and perch height selection in a guild of Libellulidae species (Odonata). Intenational Journal of Odonatology 9:235-250.
Worthen, W. B., and E. R. Patrick*. 2004. The effect of intraspecific and interspecific interactions on perch-height preferences of three odonate taxa. International Journal of Odonatology 7:529-541.
Worthen, W. B., S. Gregory*, J. Felten*, and M. J. Hutton*. 2004. Larval habitat associations of the Common Sanddragon (Progomphus obscurus Rambur) at two spatial scales (Odonata: Gomphidae). International Journal of Odonatology 7:97-109.
Worthen, W. B. 2003. Nested subset structure of larval odonate assemblages in the Enoree River (USA). International Journal of Odonatology 6:79-90.
Worthen, W. B., and D. C. Haney. 2002. Desiccation tolerance of three mycophagous Drosophila species. American Midland Naturalist 147:387-392.
Worthen, W. B. 2002. Community ecology of larval odonates in the Enoree River basin of South Carolina. Southeastern Naturalist 1:205-216.
Worthen, W. B., T. E. Blue*, D. C. Haney, and C. B. Andersen. 2001. Abundance of Boyeria vinosa larva in the Enoree River basin, USA: chemical, physical, and biological correlates. International Journal of Odonatology 4:231-241.
Worthen, W. B., C. C. Cuddy*, D. C. Haney, V. L. Turgeon, and C. B. Andersen. 2001. The effect of an industrial spill on the macrofauna of a South Carolina stream: physiological to community-level responses. Journal of Freshwater Ecology 3:467-477.
Andersen, C. B., W. B. Worthen, and B. Polkinghorn. 2001. Humanism in the environmental sciences: a reevaluation. Journal of College Science Teaching 31:202-206.
Worthen, W. B., and D. C. Haney. 1999. Temperature tolerance in three mycophagous Drosophila species: relationships with community structure. Oikos 86:113-118.
Worthen, W. B., M. T. Jones*, and R. M. Jetton*. 1998. Community structure and environmental stress: desiccation promotes nestedness in mycophagous fly communities. Oikos 81:45-54.
Rohde, K., W. B. Worthen, M. Heap, B. Hugueny, and J. Guegan. 1998. Nestedness in assemblages of metazoan ecto- and endoparasites of marine fish. Int. J. Parasitol. 28:543- 549.
Worthen, W. B. 1996. Membership patterns and nested subset analyses: Basic descriptors for community ecology. Oikos 76:417-426.
Worthen, W. B. 1996. Latitudinal variation in the development time and mass of Drosophila melanogaster: comparisons within and between populations along a latitudinal gradient. Evolution 50:2523-2529.
Worthen, W. B., and K. Rohde. 1996. Nested subset analyses of colonization-dominated communities: metazoan ectoparasites of marine fishes. Oikos 75:741-748.
Worthen, W. B., M. L. Carswell*, and K. A. Kelly*. 1996. Nested subset structure of larval mycophagous fly assemblages: Nestedness in a non-island system. Oecologia 107:257-264.
Worthen, W. B., B. R. Bloodworth*, and M. B. Hobbs*. 1995. Habitat variability in the effects of predation and microclimate on mycophagous fly communities. Ecography 18:248-258.
Worthen, W. B., S. Mayrose*, and R. G. Wilson*. 1994. Complex interactions between predation, rainfall, and microclimate: Effects on mycophagous fly communities. Oikos 69:277-286.
Worthen, W. B., M. N. Hipp*, C. T. Twardokus*, and R. Roller*. 1993. Effects of ant predation and larval density on mycophagous fly communities. Oikos 66:526-532.
Lewis*, G. P. and W. B. Worthen. 1992. Effects of predation and microenvironment on the survival of Drosophila tripunctata larvae. Oikos 64:553-559.
Worthen, W. B. and J. L. Moore*. 1991. Higher order interactions and indirect effects: A resolution using laboratory Drosophila communities. American Naturalist 138: 1092-1104.
White, D. W., W. Worthen, and E. W. Stiles. 1990. Woodlands in a post-agricultural landscape in New Jersey. Bulletin of the Torrey Botanical Club 117:256-265.
Worthen, W. B. and T. R. McGuire. 1990. Predictability of ephemeral mushrooms and implications for mycophagous fly communities. American Midland Naturalist 124:12-23.
Worthen, W. B. 1989. Effects of resource density on mycophagous fly dispersal and community structure. Oikos 54:145-153.
Worthen, W. B. 1989. Predator-mediated coexistence in laboratory communities of mycophagous Drosophila (Diptera: Drosophilidae). Ecological Entomology 14:117-126.
Worthen, W. B. 1988. Slugs (Arion spp.) facilitate mycophagous drosophilids in laboratory and field experiments. Oikos 53:161-166.
Worthen, W. B. and T. R. McGuire. 1988. A criticism of the aggregation model of coexistence: non- independent distribution of dipteran species on ephemeral resources. American Naturalist 131:453-458.
Worthen, W. B. and E. W. Stiles. 1988. Pollen-limited fruit set in isolated patches of Maianthemum canadense (Desf.) in New Jersey. Bulletin of the Torrey Botanical Club 115:299-305.
Worthen, W. B. and E. W. Stiles. 1986. Phenotypic and demographic variability among patches of Maianthemum canadense (Desf.) in central New Jersey, and the use of self-incompatibility for clone discrimination. Bulletin of the Torrey Botanical Club 113:398-405.
Worthen, W. B., A. S. Henderson, P. R. Rasmussen, and T. L. Benson (Eds.). 2009. Competition: A multidisciplinary analysis. Boston, MA: Sense.
Worthen, W. B. 2009. Dominance of the competition paradigm. IN: W. B. Worthen, A. S. Henderson, P. R. Rasmussen, and T. L. Benson (Eds.). Competition: A multidisciplinary analysis. (pp. xiii-xv). Boston, MA: Sense.
Worthen, W. B. 2009. Darwin's entangled bank: the competition paradigm in ecology. IN: W. B. Worthen, A. S. Henderson, P. R. Rasmussen, and T. L. Benson (Eds.). Competition: A multidisciplinary analysis. (pp. 17-31). Boston, MA: Sense.
ARTICLES:
Worthen, W. B. 2009. A second Odonata survey at Congaree National Park, Richland Co., SC. Argia 21: 19.
Worthen, W. B. 2009. The Darwin Legacy. Furman Magazine 52(3): 2-7.
Worthen, W. B. 2008. Fueling the Fire: point-counterpoint on offshore drilling. Counterpoint: Don't Drill. G Magazine (The Magazine of Greenville), Sept-Oct 2008 issue, pp. 63, 65.
Worthen, W. B., and C. M. Jones*. 2007. Odonata Survey of Union County, South Carolina. Argia 18: 32-33.
Bried, J., P. Hunt, and W. B. Worthen. 2007. How Often and How Long? Studying Temporal Survey Design for Adult Odonates. Argia 18:7-8
Worthen, W. B. 2005. Odonata survey of Paris Mountain State Park and Jones Gap State Park, Greenville County, South Carolina. Argia 17:27-28.
Worthen, W. B. 2003. A survey of odonates of Congaree Swamp National Monument, Richland Co., South Carolina. Argia 15:14-16.
Worthen, W. B. 2003. A Guide to the Dragonflies of Congaree Swamp National Monument.
Worthen, W. B. 2001. New dragonfly records for Sierra County, New Mexico. Argia 13:14.
Horne*, C., W. B. Worthen W. B., and K. Fravel*. 2020. The effect of habitat parameters at several scales on the abundance and diversity of larval dragonflies (Insecta: Odonata). Meeting of the South Carolina Independent Colleges and Universities, Spartanburg, SC, 20 Feb, 2020.
Horne*, C., W. B. Worthen W. B., and K. Fravel*. 2020. The effect of habitat parameters at several scales on the abundance and diversity of larval dragonflies (Insecta: Odonata). Annual Meeting of the Association of Southeastern Biologists, March 2020. (Cancelled – COVID).
Fravel*, K., W. B. Worthen, and C. Horne*. 2020. Changes in adult odonate communities along the Reedy River in Greenville Co., SC: habitat associations at several spatial scales. Annual Meeting of the Association of Southeastern Biologists, March 2020. (Cancelled – COVID).
McGovern*, P. J., and W. B. Worthen. 2020. Spatiotemporal patterns in water mite (Arrenurus spp.) parasitism of dragonflies. Annual Meeting of the Association of Southeastern Biologists, March 2020. (Cancelled – COVID).
Beck*, P., and W. B. Worthen. 2020. The effect of male maturity on intra- and interspecific aggression by the dragonfly, Erythemis simplicicollis. Annual Meeting of the Association of Southeastern Biologists, March 2020. (Cancelled – COVID).
Chamlee*, M., and W. B. Worthen. 2018. Determinants of dragonfly community structure at several spatial scales: Implications for conservation. Annual Meeting of the Association of Southeastern Biologists.
Hart*, T. M., and W. B. Worthen. 2015. The effect of water mite parasitism on fluctuating asymmetry and immune response of two dragonfly host species, Argia fumipennis and Pachydiplax longipennis. Annual Meeting of the Association of Southeastern Biologists.
Hetherington*, L. D., H. E. Clevenger*, and W. B. Worthen. 2014. Small-Scale Habitat Preferences of Ground Beetles (Coleoptera: Carabidae) and Responses to Vegetative Regrowth After Kudzu Removal. Annual Meeting of the Association of Southeastern Biologists.
Horacek*, H. J., and W. B. Worthen.2014. The Distribution of Dragonfly Larvae in a South Carolina Stream: Relationships With Sediment Type, Body Size, and the Presence of Other Larvae. Annual Meeting of the Association of Southeastern Biologists.
Turner*, L. and W. B. Worthen. 2014. Community structure of dragonfly assemblages and effects on parasitism rate: A test of the “Dilution Effect”. Annual Meeting of the South Carolina Independent Colleges and Universities.
Turner*, L. and W. B. Worthen. 2014. Community structure of dragonfly assemblages and effects on parasitism rate: A test of the “Dilution Effect”. Annual Meeting of the Association of Southeastern Biologists..
Worthen, W. B. 2013. Battling the green menace: an attempt at kudzu removal and restoration. Annual Meeting of the South Carolina Exotic Pest Plant Council, Rock Hill, SC. Sept 12, 2013.
Merriman*, D. and W. B. Worthen. 2012. Comparing biodiversity across taxonomic groups: Carabid beetle communities and local tree size and diversity. Annual Meeting of the Association of Southeastern Biologists.
Merriman*, D., A. Schulz*, and W. B. Worthen. 2012. The effects of forest structure on beetle, butterfly , and moth diversity at Furman forest. Annual Meeting of the South Carolina Independent Colleges and Universities.
DeLancey*, B. and W. B. Worthen. 2011. Changes in microhabitat selection and head dimensions between three different age classes of Desmognathus quadramaculatus. Southeastern Biology 58: 408.
Chandler*, L. and W. B. Worthen. 2011. The effects of body size and interspecific competition on perch height of libellulid dragonflies. Annual Meeting of the South Carolina Independent Colleges and Universities.
DeLancey*, B., and W. B. Worthen. 2009. Habitat selection in three species of co-occurring desmognathine salamanders in Greenville Co, SC. Southeastern Biology 55.
DeLancey*, B., and W. B. Worthen. 2009. Habitat selection in three species of co-occuring desmognathine salamanders in Greenville Co, SC. Annual Meeting of the South Carolina Academy of Science, Columbia, SC.
Worthen, W. B. 2008. The Furman Lake Restoration Project. American Association for Sustainability in Higher Education. Raleigh, NC. Nov 9-11, 2008.
Worthen, W. B. 2008. Dragonflies at Congaree: A report on a 2002 survey and plans for a 2008 survey and experiment on perch-height. Congaree National Park Research Symposium, Columbia, SC.
Jones*, C. M. and W. B. Worthen. 2006. Relationships between morphology and perch height preference in a guild of summer dragonflies. Southeastern Biology 53:217.
Gregory*, S. K., M. J. Hutton*, and W. B. Worthen. 2002. Habitat selection by larval dragonflies: correlations with sediment particle size. Southeastern Biology 49:108-109.
Haney, D. C., A. Mitchum*, W. B. Worthen, C. B. Andersen, and J. Wheeler. 2001. Effects of land use on fish diversity and abundance in the Enoree River, SC. Southeastern Biology 48:144.
Felten*, J., W. B. Worthen, and C. B. Andersen. 2001. Biological description of Enoree River (SC) tributaries: Effects of water chemistry and habitat on abundance of Progomphus obscurus. Southeastern Biology 48:109.
Mitchum, A.*, D.C. Haney, W.B. Worthen,and J. Porterfield. 2001. Fish distribution in the Enoree River, SC. Southeastern Biology 48:93.
O'Daniel*, E., L. Fikes*, W. B. Worthen, D.C. Haney, and C. B. Andersen. 2001. Comparison of diversity and abundance of larval odonate populations in rural and urban streams in the Enoree River watershed, SC. Southeastern Biology 48:110.
Blue*, T. E., W. B. Worthen, C. Cuddy*, D. C. Haney, C. B. Andersen, and S. Wheeler. 2000. The abundance of Boyeria vinosa larva (Odonata: Aeshnidae) in comparison to the physical and chemical properties of the Enoree River of South Carolina. Bulletin of the Association of Southeastern Biologists 47:101-102.
Cuddy*, C., W. B. Worthen, D. C. Haney, T. Blue*, C. B. Andersen, and S. Wheeler. 2000. Human impact on the ecology of the Upper Enoree River. Bulletin of the Association of Southeastern Biologists 47:180.
Haney, D. C., W. B. Worthen, C. B. Andersen, K. A. Sargent, and S. Wheeler. 2000. Interdisciplinary teaching using a watershed research program. Bulletin of the Association of Southeastern Biologists 47:167.
Horner*, A., A. Mitchum*, D. C. Haney, W. B. Worthen, C. B. Andersen, and S. Wheeler. 2000. Biological and chemical description of Enoree River (SC) tributaries: Effects of land use patterns on the Upper Enoree, Beaverdam Creek, and Mountain Creek. Bulletin of the Association of Southeastern Biologists 47:181.
Mitchell*, M., S. Monheim*, D. C. Haney, W. B. Worthen, C. B. Andersen, and S. Wheeler. 2000. Biological and chemical description of Enoree River (SC) tributaries II: Effects of land use patterns on Rocky Creek, Gilder Creek, and Mountain Creek. Bulletin of the Association of Southeastern Biologists 47:105.
Haney, D. C., A. Horner*, W. B. Worthen, A. Mitchum*, M. Mitchell*, C. B. Andersen, J. Wheeler. 2000. A Biological study of Enoree River (SC) tributaries: Effects of land use patterns and physicochemical measures on fish diversity and abundance. Annual Meeting of the American Society for Ichthylogy and Herpetology.
Cuddy*, C. C., D. C. Haney, W. B. Worthen, T. Blue*, A. Horner*, C. B. Andersen, J. Wheeler. 2000. Anthropogenic impact on the biology and water chemistry of the Upper Enoree River. Annual Meeting of the American Society for Ichthylogy and Herpetology.
Haney, D. C. and W. B. Worthen. 1999. Temperature tolerance in three mycophagous Drosophila species: relationships with community structure. Bulletin of the Association of Southeastern Biologists 46: 169.
Andersen, B., K. A. Sargent, W. B. Worthen, and R. Kerstetter. 1997. The Paris Mountain, SC, watershed project: a public education opportunity. American Water Resources Association, Keystone, CO. Univ. Council on Water Resources Symposium Proceedings 1997, pg. 621.
Jones*, M. T., R. H. Jetton*, and W. B. Worthen. 1997. Desiccation stress contributes to the nested subset structure of mycophagous fly communities. Bulletin of the Association of Southeastern Biologists 44:83.
Jetton*, R. H., M. T. Jones*, and W. B. Worthen. Desiccation stress contributes to the nested subset structure of mycophagous fly communities. National Counference on Undergraduate Research.
Worthen, W. B. 1996. Temperature effects on short-term responses to bi-directional selection by tropical and temperate populations of Drosophila melanogaster. Bulletin of the Association of Southeastern Biologists 43:133-134.
Worthen, W. B. 1995. The effect of temperature on response to selection. Southeastern Population Ecology and Genetics. Highlands, NC.
Carswell*, M. L. , W. B. Worthen, and K. A. Kelly*. 1995. Species-area relationships and nested-subset structure in mycophagous fly communities. Bulletin of the Association of Southeastern Biologists 42: 101.
Kelly*, K. A, , W. B. Worthen, and M. L. Carswell*. 1995. Species-area relationships and nested-subset structure in mycophagous fly communities. Bulletin of the Association of Southeastern Biologists 42: 101.
Lucas*, L. M., Mary H. Henderson*, Ron C. Michaelis, Jack Tarleton, Wade B. Worthen. 1995. Uniparental disomy in Prader-Willi syndrome. National Conference on Undergraduate Research, Union, NY.
Bloodworth*, B. R., W. B. Worthen, and M. B. Hobbs*. 1994. The effects of predation and microclimate on mycophagous flies: variation among habitats. Bulletin of the Association of Southeastern Biologists 41: 117).
Hobbs*, M. B., W. B. Worthen, and B. R. Bloodworth*. 1994. The effects of predation and microclimate on mycophagous flies: varaition among habitats. National Conference on Undergraduate Research, Kalamazoo, MI.
Worthen, W. B., B. R. Bloodworth*, and M. B. Hobbs*. 1994. Habitat variability in the effects of microclimate and ant predation on mycophagous fly communities. International Congress of Dipterology, Guelph,Ont.
Mayrose*, S., W. B. Worthen, and R. G. Wilson*. 1993. The effects of ant predation, larval density, and soil moisture on the structure of mycophagous fly communities. Bulletin of the Association of Southeastern Biologists 40:147.
Wilson*, R. G., W. B. Worthen, and S. Mayrose*. 1993. The effects of ant predatioin, larval density, and soil moisture on the survival of Drosophila tripunctata larvae. Bulletin of the Association of Southeastern Biologists 40:147.
Worthen, W. B., Sara Mayrose*, and R. Gayle Wilson*.1993.The effects of biotic and abiotic factors on larval survival and guild structure of mycophagous flies. Bulletin of the Ecolological Society of America 74:494.
Perry*, T. W., C. Prudhomme, and W. B. Worthen. 1992. Winter browse availability and use by white- tailed deer. Assoc. of Southeastern Biologists, Tuscaloosa, AL.
Worthen, W. B. 1991. Effects of predation and competition on the survival of Drosophila tripunctata larvae in a South Carolina woodland. Bulletin of the Ecological Society of America 72: 294.
Lewis*, G. P. and W. B. Worthen. 1991. Effects of predation and microclimate on survivorship of larval Drosophila. National Conference on Undergraduate Research, Pasadena, CA.
Worthen, W. B. and J. L. Moore*. 1990. Indirect effects among three mycophagous Drosophila species in laboratory communities. Bulletin of the Ecological Society of America 71:373.
Worthen, W. B. and T. R. McGuire. 1987. Community ecology on an ephemeral resource: are mycophagous fly species randomly associated, and are mushrooms really ephemeral? Bulletin of the Ecological Society of America 68:452.
Worthen, W. B. and E. W. Stiles. 1985. Pollinator limitation of fruit set in Maianthemum canadense and evidence of density dependent foraging by pollinators. Bulletin of the New Jersey Academy of Science 30: 49.
Worthen, W. B. 1988. The effects of resource distribution, facilitation and predation on the structure of mycophagous fly communities: manipulations of a model system. Ph.D. Dissertation, Rutgers, The State University of New Jersey.
Worthen, W. B. 1985. Reproductive biology of Maianthemum canadense: patch variability and interactions with the biotic environment. M.S. Thesis, Rutgers, The State University of New Jersey.
Worthen, W. B. 2009. Dragonflies at Congaree. Meeting of the Butterfly Society of the Carolinas. June 27, 2009.
Worthen, W. B. 2004. Environmental stress and community structure: patterns in terrestrial and aquatic insect communities. July 13, 2004. Berry College, Rome, GA.
Worthen, W. B. 2003. Environmental stress and community structure: patterns in larval dragonfly communities. South Carolina Association of Naturalists, Jan 15, 2003.
Worthen, W. B. 2002. Environmental stress and community structure: patterns in terrestrial and aquatic insect communities. The 15th Annual Mike Duke Memorial Seminar, The Department of Entomology, North Carolina State University.
Worthen, W. B. 2002. Environmental stress and community structure: patterns in terrestrial and aquatic insect communities. Biology Dept., University of South Carolina - Spartanburg.
Worthen, W. B. 2002. Environmental stress and community structure: patterns in terrestrial and aquatic insect communities. Graduate Program in Entomology, Clemson University, Clemson, SC.
Worthen, W. B. 2000. The physiological determinants of community structure. Berry College, Berry, GA.
Worthen, W. B. 2000. Studies in mycophagous fly ecology: from physiology to community structure. Western Carolina University, Cullowhee, NC.
Worthen, W. B. 1997. Latitudinal gradients in species diversity: why are the tropics so diverse? Dept. of Biology Newberry College, Newberry, SC.
Worthen, W. B. 1995. Nestedness and non-random community structure. Graduate Program in Zoology, University of New England, Armidale, NSW (Australia).
Worthen, W. B. 1995. The effects of habitat subdivision on biodiversity. Dept. of Biology, Anderson College, Anderson, SC.
Worthen, W. B. 1992. The effects of biotic and abiotic factors on the structure of mycophagous fly communities. Graduate Program in Entomology, Clemson University, Clemson, SC.