Step #1

VP is the combination of genetic and environmental variance. The variance from the F1 generation can be averaged with the P1 generation average variance to give the estimation of the environmental input.

(

3.1

3.9

) / 2 =

3.5 (average of the P

variance)

(

3.5

4.7

) / 2 =

4.1

Estimation of Environmental Input

VP is the combination of genetic and environmental variance. The variance from the F1 generation can be averaged with the P1 generation average variance to give the estimation of the environmental input.