Photosynthesis:
Carbon Metabolism
Readings for Chapter 10:
-
Skim pages 190-195: This portion of
the chapter tells about the C3 pathway of photosynthetic carbon reduction
(PCR) or carbon fixation.
-
Read pages 195-197: Activation and Regulation
of the C3 PCR cycle.
-
Skim pages 197-199: This portion of
the chapter tells about photorespiration.
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Read pages 199-200: Why Photorespiration?
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Skim pages 200-204: This portion of
the chapter goes over the details of the C4 pathway.
-
Read pages 204-205: Ecological Significance
of the C4 Syndrome
-
Read pages 206-207: CAM
-
Read pages 208: Regulation of C4 and CAM
-
Skim pages 208-212: Starch and Sucrose.
Goals and Objectives:
-
Be able to describe why the C3 pathway is
said to be autocatalytic and why it is considered beneficial to the plant.
-
Be able to relate the C3 pathway mechanics
to photorespiration. Explain why a proper balance of CO2
and O2 is required by the plant. In so doing, realize
the importance of controlling transpiration at the same time.
-
Be able to discuss the ecological importance
of photorespiration.
-
Be able to relate the importance of leaf structure
in the C4 plant to the function of C4 metabolism. Relate this to
photorespiration.
-
Be able to compare and contrast C3 and C4
metabolism.
-
Be able to describe the significant way in
which CAM differs from C4 metabolism.
Study Questions:
The Photosynthetic Carbon Reduction (PCR)
Cycle
-
Review the process behind C3 metabolism.
Pay particular attention to RuBisCo (pages 189-195)
Activation and Regulation of the PCR Cycle
(pages 195-197)
-
Why is it important to regulate the pool of
intermediates in PCR?
-
How do light, pH, Mg2+, and CO2
control Rubisco activity?
-
How does Rubisco activase function in controlling
Rubisco activity?
-
There are four other light activated enzymes
in PCR. What mechanism for regulation is used with these four enzymes?
Photorespiration
-
Review the process of photorespiration.
Familiarize yourself with the process, the organelles, the intermediates,
and the enzymes involved. (pages 197-199
-
Why does photorespiration occur?
-
Compare/contrast the problems that a C3 plant
would have balancing transpiration, CO2, and O2 in
a leaf.
C4 Metabolism
-
Review the processes involved in C4 metabolism
and the structure of the C4 leaf (pages 200-203)
-
Compare/contrast C3 and C4 metabolism processes
for the following (pages 204-205
-
Photorespiration
-
CO2 compensation concentration
-
Temperature Optimum
-
Quantum yield
-
Transpiration Ratio
-
Light saturation
CAM
-
Review the process of CAM (pages 206-207)
-
What are the differences/similarities between
C4 and CAM (pages 207)
-
Describe why CAM plants are well adapted to
dry environments (pages 207)
Regulation of C4 and CAM
-
How is PEPcase different in C4 and CAM plants?
-
How is PEPcase activated in the C4 plant?
-
How is activation of PEPcase accomplished
in CAM plants?